L-glutamyl-tRNA(Glu) + L-arginyl-tRNA(Arg) <=> cyclo(L-arginyl-L-glutamyl) + tRNA(Glu) + tRNA(Arg) + 2 H(+)
l-glutamyl-trna(glu) + l-arginyl-trna(arg) <=> cyclo(l-arginyl-l-glutamyl) + trna(glu) + trna(arg) + 2 h(+)