guanosine(out) + H(+)(in) => guanosine(in) + H(+)(out)
guanosine(out) + h(+)(in) => guanosine(in) + h(+)(out)
guanosine(out) + H(+)(in) => guanosine(in) + H(+)(out)
guanosine(out) + h(+)(in) => guanosine(in) + h(+)(out)