sulfate(out) + succinate(in) <=> sulfate(in) + succinate(out)
sulfate(out) + succinate(in) <=> sulfate(in) + succinate(out)
sulfate(out) + succinate(in) <=> sulfate(in) + succinate(out)
sulfate(out) + succinate(in) <=> sulfate(in) + succinate(out)