androst-4-ene-3,17-dione + AH2 + O2 <=> testololactone + A + H2O
androst-4-ene-3,17-dione + ah2 + o2 <=> testololactone + a + h2o
androst-4-ene-3,17-dione + AH2 + O2 <=> testololactone + A + H2O
androst-4-ene-3,17-dione + ah2 + o2 <=> testololactone + a + h2o