(S)-malate(out) + succinate(in) => (S)-malate(in) + succinate(out)
(s)-malate(out) + succinate(in) => (s)-malate(in) + succinate(out)
(S)-malate(out) + succinate(in) => (S)-malate(in) + succinate(out)
(s)-malate(out) + succinate(in) => (s)-malate(in) + succinate(out)