a 1-hydroxy-3-oxo steroid + A <=> a steroid 1,3-dione + AH2
a 1-hydroxy-3-oxo steroid + a <=> a steroid 1,3-dione + ah2
a 1-hydroxy-3-oxo steroid + A <=> a steroid 1,3-dione + AH2
a 1-hydroxy-3-oxo steroid + a <=> a steroid 1,3-dione + ah2