a beta-D-Gal-(1<->1')-ceramide + UDP-alpha-D-galactose <=> alpha-D-Gal-(1->4)-beta-D-Gal-(1<->1')-Cer + UDP + H(+)
a beta-d-gal-(1<->1')-ceramide + udp-alpha-d-galactose <=> alpha-d-gal-(1->4)-beta-d-gal-(1<->1')-cer + udp + h(+)
a beta-D-Gal-(1<->1')-ceramide + UDP-alpha-D-galactose <=> alpha-D-Gal-(1->4)-beta-D-Gal-(1<->1')-Cer + UDP + H(+)
a beta-d-gal-(1<->1')-ceramide + udp-alpha-d-galactose <=> alpha-d-gal-(1->4)-beta-d-gal-(1<->1')-cer + udp + h(+)