citrate(in) + 3 H(+)(in) <=> citrate(out) + 3 H(+)(out)
citrate(in) + 3 h(+)(in) <=> citrate(out) + 3 h(+)(out)
citrate(in) + 3 H(+)(in) <=> citrate(out) + 3 H(+)(out)
citrate(in) + 3 h(+)(in) <=> citrate(out) + 3 h(+)(out)