echinenone + AH2 + O2 => 4'-hydroxyechinenone + A + H2O
echinenone + ah2 + o2 => 4'-hydroxyechinenone + a + h2o
echinenone + AH2 + O2 => 4'-hydroxyechinenone + A + H2O
echinenone + ah2 + o2 => 4'-hydroxyechinenone + a + h2o