putrescine + L-glutamate + ATP = gamma-L-glutamylputrescine + ADP + phosphate + H(+)
putrescine + l-glutamate + atp = gamma-l-glutamylputrescine + adp + phosphate + h(+)
putrescine + L-glutamate + ATP = gamma-L-glutamylputrescine + ADP + phosphate + H(+)
putrescine + l-glutamate + atp = gamma-l-glutamylputrescine + adp + phosphate + h(+)