guanosine(34) in tRNA + queuine = queuosine(34) in tRNA + guanine
guanosine(34) in trna + queuine = queuosine(34) in trna + guanine
guanosine(34) in tRNA + queuine = queuosine(34) in tRNA + guanine
guanosine(34) in trna + queuine = queuosine(34) in trna + guanine