orotate(out) + a carboxylate(in) <=> orotate(in) + a carboxylate(out)
orotate(out) + a carboxylate(in) <=> orotate(in) + a carboxylate(out)
orotate(out) + a carboxylate(in) <=> orotate(in) + a carboxylate(out)
orotate(out) + a carboxylate(in) <=> orotate(in) + a carboxylate(out)