pyruvate(out) + H(+)(out) => pyruvate(in) + H(+)(in)
pyruvate(out) + h(+)(out) => pyruvate(in) + h(+)(in)
pyruvate(out) + H(+)(out) => pyruvate(in) + H(+)(in)
pyruvate(out) + h(+)(out) => pyruvate(in) + h(+)(in)