4'-hydroxyechinenone + AH2 + O2 <=> 4',4'-dihydroxyechinenone + A + H2O
4'-hydroxyechinenone + ah2 + o2 <=> 4',4'-dihydroxyechinenone + a + h2o
4'-hydroxyechinenone + AH2 + O2 <=> 4',4'-dihydroxyechinenone + A + H2O
4'-hydroxyechinenone + ah2 + o2 <=> 4',4'-dihydroxyechinenone + a + h2o