malonate(out) + succinate(in) <=> malonate(in) + succinate(out)
malonate(out) + succinate(in) <=> malonate(in) + succinate(out)
malonate(out) + succinate(in) <=> malonate(in) + succinate(out)
malonate(out) + succinate(in) <=> malonate(in) + succinate(out)