(S)-malate(in) + succinate(out) => (S)-malate(out) + succinate(in)
(s)-malate(in) + succinate(out) => (s)-malate(out) + succinate(in)
(S)-malate(in) + succinate(out) => (S)-malate(out) + succinate(in)
(s)-malate(in) + succinate(out) => (s)-malate(out) + succinate(in)