orotate(out) + a carboxylate(in) = orotate(in) + a carboxylate(out)
orotate(out) + a carboxylate(in) = orotate(in) + a carboxylate(out)
orotate(out) + a carboxylate(in) = orotate(in) + a carboxylate(out)
orotate(out) + a carboxylate(in) = orotate(in) + a carboxylate(out)