label: Phospho R-SMAD(SMAD2/3):CO-SMAD(SMAD4):FOXO3 binds FoxO3a-binding elements (Drosophila melanogaster)
label_lc: phospho r-smad(smad2/3):co-smad(smad4):foxo3 binds foxo3a-binding elements (drosophila melanogaster)