label: a (4S)-2,3-dehydroflavan-3,4-diol + succinate + CO2 + H2O + H(+) => a (2R,3S,4S)-leucoanthocyanidin + 2-oxoglutarate + O2
label_lc: a (4s)-2,3-dehydroflavan-3,4-diol + succinate + co2 + h2o + h(+) => a (2r,3s,4s)-leucoanthocyanidin + 2-oxoglutarate + o2