label: gamma-L-glutamyl-L-glutamyl-[beta-tubulin] + ADP + phosphate + H(+) => L-glutamyl-[beta-tubulin] + L-glutamate + ATP
label_lc: gamma-l-glutamyl-l-glutamyl-[beta-tubulin] + adp + phosphate + h(+) => l-glutamyl-[beta-tubulin] + l-glutamate + atp