label: gamma-L-glutamyl-L-glutamyl-[beta-tubulin] + L-glutamate + ATP = L-glutamyl-gamma-L-glutamyl-L-glutamyl-[beta-tubulin] + ADP + phosphate + H(+)
label_lc: gamma-l-glutamyl-l-glutamyl-[beta-tubulin] + l-glutamate + atp = l-glutamyl-gamma-l-glutamyl-l-glutamyl-[beta-tubulin] + adp + phosphate + h(+)