label: a beta-D-Gal-(1<->1')-ceramide + UDP-alpha-D-galactose => alpha-D-Gal-(1->4)-beta-D-Gal-(1<->1')-Cer + UDP + H(+)
label_lc: a beta-d-gal-(1<->1')-ceramide + udp-alpha-d-galactose => alpha-d-gal-(1->4)-beta-d-gal-(1<->1')-cer + udp + h(+)