preferred_name: (S)-malate(out) + succinate(in) => (S)-malate(in) + succinate(out)
preferred_name_lc: (s)-malate(out) + succinate(in) => (s)-malate(in) + succinate(out)
label: (S)-malate(out) + succinate(in) => (S)-malate(in) + succinate(out)
label_lc: (s)-malate(out) + succinate(in) => (s)-malate(in) + succinate(out)