preferred_name: orotate(in) + a carboxylate(out) => orotate(out) + a carboxylate(in)
preferred_name_lc: orotate(in) + a carboxylate(out) => orotate(out) + a carboxylate(in)
label: orotate(in) + a carboxylate(out) => orotate(out) + a carboxylate(in)
label_lc: orotate(in) + a carboxylate(out) => orotate(out) + a carboxylate(in)