preferred_name: citrate(in) + succinate(out) => citrate(out) + succinate(in)
preferred_name_lc: citrate(in) + succinate(out) => citrate(out) + succinate(in)
label: citrate(in) + succinate(out) => citrate(out) + succinate(in)
label_lc: citrate(in) + succinate(out) => citrate(out) + succinate(in)