preferred_name: Na(+)(in) + 2 H(+)(out) => Na(+)(out) + 2 H(+)(in)
preferred_name_lc: na(+)(in) + 2 h(+)(out) => na(+)(out) + 2 h(+)(in)
label: Na(+)(in) + 2 H(+)(out) => Na(+)(out) + 2 H(+)(in)
label_lc: na(+)(in) + 2 h(+)(out) => na(+)(out) + 2 h(+)(in)