preferred_name: orotate(out) + a carboxylate(in) <=> orotate(in) + a carboxylate(out)
preferred_name_lc: orotate(out) + a carboxylate(in) <=> orotate(in) + a carboxylate(out)
label: orotate(out) + a carboxylate(in) <=> orotate(in) + a carboxylate(out)
label_lc: orotate(out) + a carboxylate(in) <=> orotate(in) + a carboxylate(out)