preferred_name: malonate(in) + succinate(out) => malonate(out) + succinate(in)
preferred_name_lc: malonate(in) + succinate(out) => malonate(out) + succinate(in)
label: malonate(in) + succinate(out) => malonate(out) + succinate(in)
label_lc: malonate(in) + succinate(out) => malonate(out) + succinate(in)