preferred_name: shikimate(out) + H(+)(out) => shikimate(in) + H(+)(in)
preferred_name_lc: shikimate(out) + h(+)(out) => shikimate(in) + h(+)(in)
label: shikimate(out) + H(+)(out) => shikimate(in) + H(+)(in)
label_lc: shikimate(out) + h(+)(out) => shikimate(in) + h(+)(in)