preferred_name: succinate(out) + 2 H(+)(out) => succinate(in) + 2 H(+)(in)
preferred_name_lc: succinate(out) + 2 h(+)(out) => succinate(in) + 2 h(+)(in)
label: succinate(out) + 2 H(+)(out) => succinate(in) + 2 H(+)(in)
label_lc: succinate(out) + 2 h(+)(out) => succinate(in) + 2 h(+)(in)